Imbricatea Cavalier-Smith 2012
Cells with secreted surface silicious scales.
1.0 Euglyphida Copeland 1956, emend. Cavalier-Smith 1997
Test of organic material; most taxa with secreted silica scales held together by an organic cement.
1.1. Euglyphidae Wallich 1864, emend Lara et al. 2007
Thin, overlapping, elliptical scales; presence of specialized scales around the aperture with typical indentation.
1.1.1. Euglypha Dujardin 1841
Test elongate ovoid or pyriform; aperture with denticulate mouth pieces.
Euglypha rotunda Wailes 1911
Euglypha compressa Carter 1864
1.2. Trinematidae Hoogenraad & De Groot 1940, emend Adl et al. 2012
Test with bilateral symmetry; scales oval or round, sometimes of both types; specialized tooth-shaped scales around the aperture; aperture invaginated in some taxa.
1.2.1. Trinema Dujardin 1841
Test with large circular scales and smaller circular or ellipsoid scales; with tooth-shaped apertural scales.
1.3. Cyphoderiidae de Saedeleer 1934
Scales circular, oval or kidney-shaped, juxtaposed or imbricated; test aperture angled, some with collar.
1.3.1. Cyphoderia Schlumberger 1845
Test retort-shaped; colorless to yellow; made up of a thin chitinous membrane, covered with discs or scales; aperture terminal, oblique, circular; body does not fill the test completely; nucleus large, posterior; pseudopodia, few, long filose, simple or branched; fresh water.
1.4. Sphenoderiidae Chatelain 2013
Self-secreted scales, circular or elliptical; aperture surrounded by small round or oval scales.
1.4.1. Sphenoderia Schlumberger 1845
Test clear, circular or ovoid, covered with overlapping elliptical or nearly circular scales. Aperture circular or a narrow, oval slit at the end of a broad, clear collar covered with numerous small plates. Nucleus spherical, with few small nucleoli.
Sphenoderia lenta Schlumberger 1845
1.4. Paulinellidae De Sandeleer 1934
1.4.1. Paulinella Lauterborn 1895
Shell pyroid or broad-elliptical, not compressed, composed of siliceous, rectanguloid plates with slightly rounded, scarcely overlapping borders, arranged in circumferential longitudinal rows; in apertural view the plates form a counter-clockwise spiral, with one pentagonal scale at the aboral pole. Pseudostome small. Plasmabody with usually two sausage-like endosymbionts of cyanobacterial origin. Filopodia, usually fast moving.
Paulinella chromatophora Lauterborn 1895
2.0. Placoperla Cavalier-Smith 2012
Imbricate cercozoa that secrete a test of mineralized spherical pearls or silicified plate scales that are typically double tiered with a perforate upper tier.
2.1. Thaumatomonadida Shirkina1987
Heterotrophic usually gliding cells that may swim also; with flattened cell body and with two heterodynamic cilia inserting subapically and/or ventrally; some unikont; with extrusomes; filopodia produced subapically or from ventral groove; cysts; multinucleate and multiciliate stages known.
2.1.1. Thaumatomonas de Saedeleer 1931
Bicilated, heterotrophic amoeboflagellates with silicious body scales.
Euglyphida genera incertae sedis
Pareuglypha Penard 1902 (incertae sedis)
Test ovoid, tapered towards a circular aperture. With a scaly spine of variable length.
Pareuglypha reticulata Penard 1902